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Beatrice “Beezus” Quimby (Selena Gomez) is a girl who has to endure the antics of her imaginative younger sister, Ramona Quimby (Joey King). She is given the nickname Beezus by Ramona because the latter could not say Beatrice when she first learned to talk. Beezus is frequently annoyed or embarrassed by Ramona’s behavior throughout the story. In one case, Ramona scribbles in a library book that Beezus then has to purchase. In another, Ramona ruins Beezus’ checker game with Beezus’ friend Henry Huggins  and locks Ribsy, Henry’s dog, in the bathroom. Henry leaves and Beezus becomes angry. Ramona also invites numerous friends over without her mother’s permission because she wants to have a party. Later in the story, she ruins Beezus’ birthday cakes. She destroys one birthday cake and when Mrs. Quimby endeavours to bake another, Ramona manages to spoil that as well.

Beezus often feels guilty because she knows she is supposed to love Ramona, but has a hard time with that. While she tries to read to her and care for her, Beezus often gets mad about the things Ramona does. Frustrated, Beezus asks their mother if she was like Ramona when she was younger. Their mother replies that yes, she was different in some ways, such as being quieter. This pleases Beezus, who dislikes Ramona’s tendency to be very noisy. Then Mrs. Quimby explains how she and her sister, Beezus and Ramona’s Aunt Beatrice, were also very different as children. Beezus begins to think about her mom and aunt, and how much they love each other, and realizes that Ramona is not such a bad little sister after all.